Mr. Greedy, 33, was the backbone of the zoo’s Penguin Coast, an attraction that features the largest colony of endangered African penguins in North America. The cause of death was euthanasia due ...

Rest in peace, Mr. Greedy. The 33-year-old African penguin, who made a “tremendous contribution” to the survival of his endangered species by fathering 230 chicks, was euthanized for age ...

Mr. Greedy is survived by his lifelong mate, African penguin No. 832, known as Mrs. Greedy. She is now the oldest penguin in the Maryland Zoo’s colony at 33.